DNA-directed RNA polymerase RpoA/D/Rpb3-type
Large ribosomal subunit protein uL30-like ferredoxin-like fold
Small ribosomal subunit protein uS4 N-terminal
Isoglutamyl lysine isopeptide (Gln-Lys) (interchain with K-? in acceptor proteins)
Reversibly protonated during proton transport